Center for Ecological Studies

PAN Kaiwen Research Group

PAN Kaiwen Research Group

Update time: 09/02/2019

PI: Dr. Kaiwen Pan, professor, Doctoral tutor, group leader 

Chief scientist of Ecological Processes and Biodiversity Research Group

Chengdu Institute of Biology 
Chinese Academy of Sciences
Chengdu, Sichuan, 610041






1987. 9 – 1991. 7, B. Sc. Degree in forestry from the Department of Forestry, Sichuan Agricultural University, Yaan, Sichuan Province, P. R. China; 

1991. 9 – 1994. 7, M. Sc. Degree in Silviculture from the Faculty of Forestry & Horticulture in Sichuan Agricultural University, Sichuan Province, P. R. China; 

1994. 9 -1997. 6,  PhD degree in ecology science (plant community structure, function and dynamics) from Shenyang Institute of Applied Ecology, the Chinese Academy of Sciences, Shenyang, Liaoning Province, P. R. China 



1997.7 - 1999. 11, Assistant researcher, Center for Botanic Studies, Chengdu Institute of Biology (CIB), the Chinese Academy of Sciences (CAS), P. R. China; 

1999.12 - 2002. 7, Associate professor, Center for Studies on Restoration of Degradation Ecosystem, CIB, CAS, P. R. China; 

2002. 8 – present, Professor, Center for Ecological Studies, CIB, CAS, P. R. China; 

2006. 6 – 2014, Vice-head and Professor in Center for Ecological Studies, CIB, P. R. China;

2014 – 2019, Principal scientist in Ecological Processes and Biodiversity Research, head of Center for Ecological Studies, CIB, P. R. China.

2019 - present, Professor Biodiversity and Ecosystem Services, CIB, P. R. China. 


Involvement in Editorial board member of group leader

Ecology and Environmental Sciences

Agricultural Science

Frontiers in Environmental Science, section Agroecology and Land Use Systems

Journal of Chinese Urban Forestry


Involvement in professional society of group leader

Deputy director of Professional Committee of Allelopathy, CSPP

The Secretary General of the Ecological Society of Sichuan Province

Deputy director of Professional Committee of Ecology, Environmental Science Society of Sichuan Province

Executive director of the Geographical Society of Sichuan Province

Vice President of Sichuan Ecotourism Association

Member of a council of Ecological Society of China

HONORS & AWARDS of group leader

2001, Silver Medal for Science & Technology Award from the People’s Government of Sichuan Province;

2006, Silver Medal for Science & Technology Award from the People’s Government of Sichuan Province

2007, Award for Expert with Outstanding Achievement from the CPC Sichuan Province Committee and the People’s Government of Sichuan Province; 

2012, Award for Outstanding Environmental Science and Technology Worker from the Ninth Chinese Environmental Science Society;

2014, Academic Technology Leaders in Sichuan Province granted by People’s Government of Sichuan Province.

2018, Advanced Individuals in the Construction of Ecological Barrier in the Upper Reaches of the Yangtze River in Sichuan Province by People’s Government of Sichuan Province.

2019Received a special government subsidy from the State Council.



2003. 9 - 2004. 9, Visiting professor affiliated with Griffith University, Australia 

2005. 8 – 2005. 10, Senior scholar in Yokohama National University, Japan 

Prof. PAN Kaiwen attended international conferences and visited relevant laboratories in Australia, Japan, USA, Denmark, and Czech Republic, etc. He had established the effective linkages of relevant collaborative researches with colleagues in USA, Japan, Australia, Czech Republic, Spain and Poland. 


Other members in the Group


Dr. Zhang Linassociate professor. He is interested in restoration ecology and soil ecology.                                                 

Dr. Sun Xiaoming, Research assistant. She is interested in soil fauna ecology and forest ecology.

Dr. Xiong Qinli, Research assistant. He is interested in landscape ecology and geographic information system.

Mr. Wu XiaogangResearch probationer. He is interested in vegetation ecology.



Mr. Li ZilongDoctoral student. He is interested in plant physioecology, especially in responses of plants to stressed drought and rainfall. Graduation.

 Mr. Wang SizhongDoctoral student. He is interested in soil fauna ecology, especially in responses of soil macrofauna to stressed drought and rainfall. Graduation.                                              

Mr. Akash Tariq from PakistanDoctoral student. He is interested in plant physioecology, especially in responses of tree species to stressed drought and rainfall. Graduation.

Mr. Olatunji Abiodun Oluanya from NigeriaDoctoral student. He is interested in soil ecology, especially in soil food webs. Graduation.

Mr. Song DagangDoctoral student. He is interested in plant community ecology. Graduation.

Mr. Zhang AipingGraduate student. He is interested in plant community ecology. Graduation.                                                     

Miss Tan XueGraduate student. She is interested in plant community ecology. Graduation.

Miss Huang DanGraduate student. She is interested in soil ecology. Graduation.

Miss Li NingningGraduate student. She is interested in soil ecology. Graduation.

Mr. Mohammed A. Dakhil from EgyptDoctoral student. He is interested in geobotany, especially plant species migration and diffusion.

Mr. Bikram Pandey from Nepal, Doctoral student. He is interested in plant functional biodiversity.


Mr. META F. J. S. from South Sudan, Doctoral student. He is interested in soil carbon and nitrogen dynamic.

Miss Hongyang Zhou, Doctoral student. He is interested in soil food web.


 Mr. Zebene Tadesse Negesse from Ethiopia, Doctoral student. He is interested in soil carbon, nitrogen dynamic and soil fauna.

Miss Zhang FenyingGraduate student. She is interested in plant community ecology.

Miss Zhang MengGraduate student. She is interested in plant community ecology.

Mr. Tang TianwenGraduate student. She is interested in plant community ecology.


Research field


It includes the dynamics of the important functional groups of macrofauna, nematodes, collembola and mites, soil food web and relative to environmental factors, and its spatial distribution pattern in large scale. We are also interested in analyzing the relationships between the underground nutrients and the soil food web, the interaction effects between the underground food webs and the aboveground plant functional groups in the context of human activities and climate change, etc. in mountainous regions of Southwestern China.


This include studies on the southwestern mountain forests, agroforestry systems and burned forest area, with the aim to explore the cycles of carbon and nitrogen, phosphorus and other elements of the forest soil and its responses to global change and human control, and as well as biological mechanism of soil animal, microorganism and enzyme. Some attentions will be paid to the studies of plant ecophysiology, forest management, crop cultivation, water and fertilizer management, soil and water conservation for improving comprehensive benefits of ecosystem services.


On the basis of investigating typical plant communities in southwestern mountainous regions, the remote sensing and geographic information system and other methods, combined with statistical methods will be used to study the spatial distribution patterns of typical plant communities, rare and endangered plants and important forest ecosystems. Its formation mechanism will be discussed on the basis of previous studies in the fields of palynology, paleoclimate, plant systemic evolution. Hot spot area will thus be identified for providing suggestions of protection, control measures and policy in this region.


There have being some ecological wound caused by important construction projects such as hydropower station, road and mining in southwestern mountain region, dry valley, and high altitude fragile area. To obtain technical system for the ecological recovery of these anthropogenic wound, suitable plant species will be screened. Soil water and fertilizer managements, plant community design as well as reconstruction studied will be our main focus.


The knowledge of botany, ecology, natural geography, landscape ecology, geographical information system will be used to study ecosystem assessment, ecological civilization construction planning, natural protected area planning, ecological protection zones delineation, ecological service function evaluation, ecological security pattern optimization, and ecological compensation. This studies will provide the basis for national and regional ecological civilization construction.

We warmly welcome individuals with good academic background in natural geography, botany, ecology, forestry, soil, biology, and as well as remote sensing and geographic information systems to join our research team for pursuing a doctor's degree or cooperative researches. University of Chinese Academy of Sciences may provide fellowships including CAS-TWAS President's Fellowship Programme for doctoral candidates, Chinese Government Scholarship—Chinese University Program for graduate study at UCASScholarship for Students from Hong Kong, Macao & Taiwan and UCAS Scholarship for International Students. The Research Fund for International Young Scientists (the Fund) by the National Natural Science Foundation of China (NSFC) also supports foreign young scientists to conduct basic research in mainland China with the aim to promote sustainable academic collaboration and exchanges between Chinese and foreign young scientists. The Chinese Academy of Sciences offers various international fellowships to support highly-qualified international scientists and postdoctoral scholars to work and study in my institute.

Selected Publication

1.     Johan van den Hoogen*, Stefan GeisenDevin RouthHoward FerrisWalter TraunspurgerDavid A. WardleRon G. M. de GoedeByron J. AdamsWasim AhmadWalter S. AndriuzziRichard D. BardgettMichael BonkowskiRaquel Campos-HerreraJuvenil E. CaresTancredi CarusoLarissa de Brito CaixetaXiaoyun ChenSofia R. CostaRachel CreamerJosé Mauro da Cunha CastroMarie DamDjibril DjigalMiguel Escuer, Bryan S. GriffithsCarmen GutiérrezKarin HohbergDaria KalinkinaPaul KardolAlan KergunteuilGerard KorthalsValentyna KrashevskaAlexey A. KudrinQi LiWenju LiangMatthew MagiltonMariette MaraisJosé Antonio Rodríguez MartínElizaveta MatveevaEl Hassan MayadChristian MulderPeter MullinRoy NeilsonT. A. Duong NguyenUffe N. NielsenHiroaki OkadaJuan Emilio Palomares RiusKaiwen PanVlada PenevaLo?c PellissierJulio Carlos Pereira da SilvaCamille PitteloudThomas O. PowersKirsten PowersCasper W. QuistSergio RasmannSara Sánchez MorenoStefan ScheuHeikki Set?l?Anna SushchukAlexei V. TiunovJean Trap, Wim van der PuttenMette Vesterg?rdCecile VillenaveLieven WaeyenbergeDiana H. WallRutger WilschutDaniel G. WrightJiue-in Yang & Thomas Ward Crowther*. Soil nematode abundance and functional group composition at a global scale. Nature  2019572194–198

2.     Mohammed A. Dakhil, QinliXiong, Emad A.Farahat, LinZhang, Kaiwen Pan*, Bikram Pandey, Olusanya Abiodun Olatunji, AkashTariq, Xiaogang Wu, Aiping Zhang, Xue Tan, Dan Huang. Past and future climatic indicators for distribution patterns and conservation planning of temperate coniferous forests in southwestern China. Ecological Indicators, 2019, 107, December, 105559.

3.     Sizhong Wang, Olusanya Abiodun Olatunji, Cao Guo, Lin Zhang*, Xiaoming Sun, Akash Tariq, Xiaogang Wu, Kaiwen Pan*, Zilong Li, Feng Sun & Dagang Song. Response of the soil macrofauna abundance and community structure to drought stress under agroforestry system in southeastern Qinghai-Tibet Plateau. Archives of Agronomy and Soil Science,

4.     Akash Tariq, Kaiwen Pan, Olusanya Abiodun Olatunji, Corina Graciano, Ningning Li, Zilong Li, Dagang Song, Feng Sun, Meta Francis Justine, Dan Huang, Shanxing Gong, Bikram Pandey, Muhammad Idrees, Mohammed A. Dakhil. Role of nitrogen supplementation in alleviating droughtassociated growth and metabolic impairments in Phoebe zhennan seedlings. Journal of Plant Nutrition and Soil Science, First published: 29 April 2019,

5.     Feng Sun, Kaiwen Pan, Olusanya Abiodun Olatunji, Zilong Li, Wenkai Chen, Aiping Zhang, Dagang Song, Xiaoming Sun, Dan Huang, Xue Tan. Specific legumes allay drought effects on soil microbial food web activities of the focal species in agroecosystem. Plant and Soil, April 2019, Volume 437, Issue 1–2, pp 455–471, Plant and Soil (2019) 437: 455.

6.     Yang Xiao?, Qinli Xiong*,? and Kaiwen Pan*. What Is Left for Our Next Generation? Integrating Ecosystem Services into Regional Policy Planning in the Three Gorges Reservoir Area of China. Sustainability 2019, 11(1), 3;

7.     Olusanya Abiodun OlatunjiShanxing GongAkash TariqKaiwen Pan*Xiaoming SunWenkai ChenLin ZhangMohammed A. DakhilDan HuangXue Tan. The effect of phosphorus addition, soil moisture, and plant type on soil nematode abundance and community composition. Journal of Soils and Sediments, (2018).

8.     Akash Tariq, Kaiwen Pan*, Olusanya A Olatunji, Corina Graciano, Zilong Li, Ningning Li, Dagang Song, Feng Sun, Xiaogang Wu, Mohammed A. Dakhil, Xiaoming Sun, Lin Zhang. Impact of phosphorus application on drought resistant responses of Eucalyptus grandis seedlings. Physiologia Plantarum, First published: 09 November 2018,

9.     Olusanya Abiodun Olatunji, Kaiwen Pan*, Akash Tariq, Lin Zhang, Xiaogang Wu, Xiaoming Sun, Hongyan Luo, Dagang Song, Ningning Li. The responses of soil microbial community and enzyme activities of Phoebe zhennan cultivated under different soil moisture conditions to phosphorus addition. iForest 2018, 11: 751-756. - doi: 10.3832/ifor2725-011

10.   Li ZPan K*et al. Legume plants may facilitate Zanthoxylum bungeanum tolerance to extreme rainfall, Scientific Reports, 2018volume 8, Article number: 15996

11.   Qinli Xiong,Yajuan Shi, Yonglong Lu, Kaiwen Pan, Mohammed A.Dakhi, Lin Zhang, Yang Xiao. Sublethal or not? Responses of multiple biomarkers in Daphnia magna to single and joint effects of BDE-47 and BDE-209. Ecotoxicology and Environmental Safety, Volume 164, 30 November 2018, Pages 164-171,

12.   Dagang SongKaiwen Pan*, Aiping ZhangXiaogang WuAkash TariqWenkai ChenZilong LiFeng SunXiaoming SunOlusanya Abiodun Olatunji & Lin Zhang. Song D, Pan K, Zhang A, et al. Optimization of growth and production parameters of walnut (Juglans regia) saplings with response surface methodology. Scientific Reports, 2018, 8(1):9992. Published: 03 July 2018. DOI: 10.1038/s41598-018-28345-6

13.   Olusanya A. Olatunji ,Hongyan Luo, Kaiwen Pan*, Akash Tariq, Xiaoming Sun, Wenkai Chen, Xiaogang Wu, Lin Zhang, Qinli Xiong, Zilong Li, Dagang Song, Aiping Zhang, Feng Sun. Influence of phosphorus application and water deficit on the soil microbiota of N2fixing and nonNfixing tree. Ecosphere, 2018 , 9 (6).First published: 13 June 2018. DOI: 10.1002/ecs2.2276.

14.   Sizhong Wang, Kaiwen Pan*, Akash Tariq, Lin Zhang, Xiaoming Sun, Zilong Li, Feng Sun, Qinli Xiong, Dagang Song, Olusanya Abiodun Olatunji. Combined effects of cropping types and simulated extreme precipitation on the community composition and diversity of soil macrofauna in the eastern Qinghai-Tibet Plateau. Journal of Soils and Sediments. 2018:1–13. First Online: 19 April 2018. Journal of Soils and Sediments (2018).

15.   Akash TariqKaiwen Pan*, Olusanya Abiodun OlatunjiCorina GracianoZilong LiFeng SunLin ZhangXiaogang WuWenkai ChenDagang SongDan HuangTan Xue & Aiping Zhang. Phosphorous fertilization alleviates drought effects on Alnus cremastogyne by regulating its antioxidant and osmotic potential. Scientific Reports, 2018, 8(1):5644.Article number: 5644 (2018). DOI: 10.1038/s41598-018-24038-2APR 4 2018

16.   Sun F, Pan KW*, Li ZL, Wang SZ, Tariq A, Olatunji OA, Sun XM, Zhang L, Shi W*Wu XG. Soybean supplementation increases the resilience of microbial and nematode communities in soil toextreme rainfall in an agroforestry system. Science of the Total Environment, 2018, 626: 776-784.

17.   Tariq A, Pan K*, Olatunji OA, Graciano C, Li Z, Sun F, Sun X, Song D, Chen W, Zhang A, Wu X, Zhang L, Mingrui D, Xiong Q and Liu C. (2017). Phosphorous Application Improves Drought Tolerance of Phoebe zhennan. Frontiers in Plant Science, 2017,8(x):1561. doi: 10.3389/fpls.2017.01561 SEP 13 2017

18.   Tariq A, Sadia S, Pan K, Ullah I, Mussarat S, Sun F, Abiodun OO, Batbaatar A, Li Z, Song D, Xiong Q, Ullah R, Khan S, Basnet BB, Kumar B, Islam R, Adnan M. (2017). A systematic review on ethnomedicines of anticancer plants. Phytotherapy Research, 2017, 31(2): 202-264. DOI: 10.1002/ptr.5751 FEB 2017

19.   Liu C, Wang Y*, Pan K, Wang Q, Liang J, Jin Y, Tariq A. (2017). The synergistic responses of different photoprotective pathways in Dwarf Bamboo (Fragesia rufa) to drought and subsequent rewatering. Frontiers in Plant Science, 2017, 8(x):849. doi: 10.3389/fpls.2017.00489 , 04 April 2017

20.   Liu CG*, Wang QW, Jin YQ, Pan KW, Wang YJ. 2017. Photoprotective and antioxidative mechanisms against oxidative damage in Fargesia rufa subjected to drought and salinity. Functional Plant Biology, 44: 302–311.

21.   Chenggang Liu, Yanjie Wang, Yanqiang Jin, Kaiwen Pan, Xingmei Zhou, Na Li. Photoprotection regulated by phosphorus application can improve photosynthetic performance and alleviate oxidative damage indwarf bamboo subjected to water stress. Plant Physiology and Biochemistry, 118: 88-97, September 2017, online 8 June 2017

22.   Song D, Pan K*, Tariq A*, Sun F, Li Z, Sun X, Zhang L, Abiodun O and WU X (2017). Large-scale patterns of distribution and diversity of terrestrial nematodes. Applied Soil Ecology. 2017, 114:161-169. DOI: 10.1016/j.apsoil.2017.02.013JUN 2017

23.   Xiong Q, Xiao Y, Ouyang Z, Pan K, Zhang L, He X,  Zheng H, Sun X, Wu X, Tariq A, Lingjuan L. (2017). Bright side? The impacts of Three Gorges Reservoir on local ecological service of soil conservation in southwestern China. Environmental Earth Sciences2017, 76(8):323. DOI: 10.1007/s12665-017-6588-7

24.   Tariq A, Adnan M, Amber R, Pan K, Mussarat S, Shinwari ZK. (2016). Ethnomedicines and anti-parasitic activities of Pakistani medicinal plants against Plasmodia and Leishmania parasites. Annals of Clinical Microbiology and Antimicrobial2016, 15(1):52. DOI: 10.1186/s12941-016-0170-0SEP 20 2016

25.    Sun F, Pan K*, Tariq A, Zhang L, Sun X*, Li Z, Wang S, Xiong Q, Song D, Olatunji OA. (2016). The response of the soil microbial food web to extreme rainfall under different plant systems. Scientific Reports. 2016,6, 37662; doi: 10.1038/srep37662  NOV 22 2016

26.   Sun F, Tariq A, Chen H, He Q, Guan Y, Pan K, Chen S, Li J, Zhao C, Wang H, Gu Y. (2016). Effect of nitrogen and phosphorus application on agricultural soil food webs, Archives of Agronomy and Soil Science, 2016 , 63 (8) :1176-1186. DOI: 10.1080/03650340.2016.1266483 ,出版2017

27.   Song D, Pan K*, Tariq A*, Azizullah A, Sun F, Li Z, Xiong Q. (2016). Adsorptive removal of toxic chromium from waste-water using wheat straw and Eupatorium adenophorum. PLoS ONE, 2016,11(12): e0167037. DOI: 10.1371/journal.pone.0167037DEC 2 2016

28.   Xiong QL, Pan KW*, Zhang L*, Wang YJ, Li W., He XJ, Luo HY. Warming and nitrogen deposition are interactive in shaping surface soil microbial communities near the alpine timberline zone on the eastern Qinghai-Tibet Plateau, southwestern China. Applied Soil Ecology, 2016, 101: 72-83. DOI: 10.1016/j.apsoil.2016.01.011MAY 2016

29.   Zhang L, Liu LJ, Pan KW*, Li W, Wang YJ, Deng MR. Post-wildfire soil and plant foliar nutrient ratios and soil fungi:bacterial ratios in alpine meadows on the southeastern Qinghai-Tibet Plateau. International Journal of Wildland Fire, 2015, 247: 933-939. DOI: 10.1071/WF14147

30.   Liu CG, Wang YJ*, Pan KW, Jin YQ, Li W, Zhang L. Effects of phosphorus application on photosynthetic carbon and nitrogen metabolism, water use efficiency and growth of dwarf bamboo (Fargesia rufa) subjected to water deficit. Plant Physiology and Biochemistry, 2015, 96:20-28. DOI: 10.1016/j.plaphy.2015.07.018NOV 2015

31.   Liu CG, Wang YJ*, Pan KW*, Jin YQ, Liang J, Li W, Zhang L. Photosynthetic carbon and nitrogen metabolism and the relationship between their metabolites and lipid peroxidation in dwarf bamboo (Fargesia rufa Yi) during drought and subsequent recovery. Trees-Structure and Function, 2015, 296:1633-1647. DOI: 10.1007/s00468-015-1241-0. Article (PDF Available) inTrees 29(6):1-15 · June 2015DEC 2015

32.   Pan KW*Gong PMWang JCWangYJLiu CGLi WZhang L. Applications of nitrate and ammonium fertilizers alter soil nematode food webs in a continuous cucumber cropping system in Southwestern Sichuan, China. Eurasian J Soil Sci 2015, 4 (4): 287 – 300. DOI: 10.18393/ejss.2015.4.287-300. Article (PDF Available) 

33.   Li W, Pan K.W.*, Wu N., Wang J.C., Wang Y.J., Zhang L. Effect of litter type on soil microbial parameters and dissolved organic carbon in a laboratory microcosm experiment. Plant Soil Environ201460(4):170-176. DOI: 10.17221/39/2014-PSE. ArticleinPlant Soil and Environment 60(4):170-176 · April 2014.

34.   Liu C.G., Wang Y.J.*, Pan K. W., Zhu T.T., Li W., Zhang L. Carbon and nitrogen metabolism in leaves and roots of dwarf bamboo (Fargesia denudata Yi) subjected to drought for two consecutive years during sprouting period. J Plant Growth Regul (2014) 33(2):243-255. DOI:10.1007/s00344-013-9367-z. Article (PDF Available) inJournal of Plant Growth Regulation 33(2):243–255 · April 2014, JUN 2014

35.   Liu C.G., Wang Y.J.*, Pan K. W.*, Zhu T.T., Li W., Zhang L. Responses of the Antioxidant Defense System to Drought Stress in the Leaves of Fargesia denudata Seedlings, the Staple Food of the Giant Panda. Russ J Plant Physiol. 2014, 61(3):374-383. DOI: 10.7868/S0015330314020080. Article · January 2014

36.   Shen X.Y., Liu C.G., Pan K.W*. Reproductive Biological Characteristics of Dendrobium Species. Reproductive biology of plants. Edited by K . R . ShivannaCRC Press 2014P 195–217Print ISBN: 978-1-4822-0132-1eBook ISBN: 978-1-4822-0133-8DOI: 10.1201/b16535-11. Chapter (PDF Available)  · January 2014

37.   Wang JinchuangChen JuanPan Kaiwen. Effect of exogenous abscisic acid on the level of antioxidants in Atractylodes macrocephala Koidz under lead stress. Environmental science and pollution research, 2013, 20(3):1441-1449, DOI: 10.1007/s11356-012-1048-0

38.   Han C.M., Li C.L., Ye S.P., Wang H., Pan K.W.*, Wu N., Wang Y.J., Li W., Zhang L. Autotoxic effects of aqueous extracts of ginger on growth of ginger seeding and on antioxidant enzymes, membrane permeability and lipid peroxidation in leaves. Allelopathy Journal (2012) 30 (2): 259-270. October 2012.

39.   Pang S.M., Pan K.W.*, Wang Y.J.*, Li W., Zhang L., Chen Q.B. Floral morphology and reproductive biology of Dendrobium jiajiangense (Orchidaceae) in Mt. Fotang, southwestern China. Flora (2012) 207(6): 469-474. DOI: 10.1016/j.flora.2012.02.008.· June 2012

40.   Cao G.X., Xue L., Li Y.,Pan K.W. The relative importance of architecture and resource competition in allocation to pollen and ovule number within inflorescences of Hosta ventricosa varies with the resource pools. Annals of Botany ,2011, 107(8): 1413-1419. DOI: 10.1093/aob/mcr085.· June 2011

41.   Pan K.W.*, Xu Z.H., et al. Application of (NH4)2SO4 to study N dynamics in hoop pine plantation and adjacent native forest of suntropical Australia: the effects of injection depth and litter addition. Journal of Soils and Sediments, 2009, 9(6): 515-525. DOI: 10.1007/s11368-009-0131-8.  December 2009

42.   Li H.Y., Pan K.W.*, Liu Q., Wang J. C. Effect of enhanced ultraviolet-B on allelopathic potential of Zanthoxylum bungeanum. Scientia Horticulturae ,2009, 119(3): 310-314. DOI: 10.1016/j.scienta.2008.08.010.  February 2009

43.   Li W., Pan K.W.*, Wu N., Wang J.C., Han C.M., Liang X.L. Effects of mixing pine and broadleaved tree/shrub litter on decomposition and N dynamics in laboratory microcosms. Ecological Research ,2009, 24(4): 761-769. DOI: 10.1007/s11284-008-0546-5.· July 2009

44.   Wang, J.C., Wu Y. , Wang Q., Peng Y.L., Pan K.W, Luo P., Wu N. Allelopathic effects of Jatropha curcas on marigold (Tagetes erecta L.). Allelopathy Journal ,2009, 24(1): 123-130. July 2009.

45.   Pan K.W.*, Xu Z.H., Blumfield T., Totua S., Lu M.X. In situ mineral N dynamics and fate of added 15NH4+ in hoop pine plantation and adjacent native forest in subtropical Australia. Journal of Soils and Sediments (2008) 8: 398-405. DOI 10.1007/s11368-008-0037-x.

46.   Han C.M., Pan K.W.*, Wu N., Wang J.C., Li W. Allelopathic effect of ginger on seed germination and seedling growth of soybean and chive. Scientia Horticulturae (2008) 116(3): 330-336. DOI: 10.1016/j.scienta.2008.01.005.  May 2008

47.   Pan K.W., Wang J.C.*, Lv K., Song L. Effects of leaf leachates of Zanthoxylum bungeanum on soil enzymes, chemical properties and its own seedlings growth. Allelopathy Journal ,2008, 22(1):153-166. July 2008.

48.   Sun S.C. *, Bao W.K., Pan K.W., Gao X.M. Restoring mountain slope forests by strip-cutting shrubs in the upper reaches of Minjiang River, Sichuan, China: a biodiversity-oriented silviculture experiment. Chinese Forestry Science and Technology, 2004, 3(1): 7-16

49.   Pan K.W.* ,Zhang yongmei,chen Qingheng,Liu Zhaoguang.. TAPPSC: An effective method of soil and water conservation during the process of reforestation? Proceedings of 12th international soil conservation organization conference, Vol III (Jiao Juren et al edition). Tsinghua University Press, 2002, p81-84.

50.   Pan K.W. et al. Restoration and rehabilitation of Degraded Forest Ecosystem on the eastern Qinghai-Tibetan Plateau, China. Proceedings of the IUFRO Conference on Restoration of Boreal and Temperate Forest, Documenting Forest Research Knowledge and Practices in Boreal and Temperate Ecosystems (Emile S. Gardiner and Lynne J. Breland edited, ISSN 1600-8790). 2002. p165-166. Danish Center for Forest, Landscape and Planning

51.   Pan KW, Zhang L, Shao YH, Fu SL*. Special observational network of soil fauna diversity in China: revealing the mystery of soils. Biodiversity Science. 2016, doi: 10.17520/biods.2016019.

52.   Li LJ, Xiong QL, Pan KW*, Zhang L*. The responses of soil protozoan communities to Quercus aquifolioides: recovery after cutting and growing season dynamics. Biodiversity Science. 2015, 23 (6): 793–801

53.   Song DG, Pan KW*. Advances in researches on ecological benefit evaluation of the Grain for Green Project in China. Journal of Sichuan Forestry Science and Technology, 2015, 36(3):45-49.

54.   Zhu TT, Li W*, Liu CG, Pan KW*. Effect of Dendrocalamus latiflorus and Eucalyptus grandis plantations on soil nematode community in Sichuan Basin. Chinese Journal of Ecology. 2014, 33(4):1034-1041

55.   Li Q, Jin GX, Deng H, Yang L, Pan KW*, Jiang Y. The Effect of 2008 Wenchuan Earthquake on staple bamboos and the habitat for Giant Pandas in Baihe Nature Reserve in Sichuan. Journal of Sichuan Forestry Science and Technology, 2014, 35(5):80-83

56.   Huang Q, Zhu SH, Li LH, Xiong Y, Shen XY, Pan KW*. A Study of insect pests damaging tree trunks of Zanthoxylum bungeanum Plantation in Aba Zang-Qiang Autonomous Prefecture. Journal of Sichuan Forestry Science and Technology, 2013, 34(6):56-59

57.   Han CM, Li CLYe SPPan KW*Wu NLi W. Comparative study on the quality and economic benefit of ginger under different cultivated patterns. Northern Horticulture2013, 22:41-43

58.   Zhang MX, Ji M, Li W*, Liu HC, Wang YJ, Zhang L, Pan KW*. Effect of land use patterns on soil aggregate stability and aggregate-associated organic carbon. Chin J Appl Environ Biol, 2013, 19(4):598-604

59.   Han CM, Li CLYe SPPan KW*Wu NLi W. Effects of different cultivation patterns on morphological index and rhizome yields of Ginger. Northern Horticulture, 2013, (12):18-20

60.   Ma QX, Li W*, Pan KW*, Zhang L, Wang YJ. Effect of continuous glucose addition on soil N transformation of the Pinus tabulaeformis and Cercidiphyllum japonicum plantations. Chin J Appl Environ Biol, 2013, 19(3):426-433

61.   Ji M, Liu HC, Li WPan KW*, et al. The present situation and strategies of Grain for Green Project of Chengdu City. Journal of Sichuan Forestry Science and Technology, 2013, 34(2):91-94

62.   Liu CG, Huang Q, Deng MR, Shen XY, Liu LJ, Zhang L, Li W, Pan KW*. Energy, material and value flows in the different systems of Chinese prickly ash-crop intercropping in the upper reaches of the Minjiang river. Ecology and Environmental Sciences, 2013, 22(1):116-122

63.   Han C MLi C LYe S PPan KW*, et al. Effects of ginger aqueous extract on soil enzyme activitymicrobial community structure and soil nutrient content in the rhizosphere soil of ginger seedlings. Acta Ecologica Sinica, 2012, 32( 2):0489-0498

64.   Liu DH, Liu YH, Xiong H, Xu FX, Pan KW, et al. Research on the harmfulness, prevention and control of main agricultural weather disaster in Southwest China. Chinese Journal of Agrometeorology. 2011, 32: 208

65.   212

66.   Li K, Zeng XB, Li ZB, Pan KW*, et al. Soil and litter water-holding capacity of Dendrocalamus latiflorus plantations on different slope positions after converting cropland to forest in Chengdu. Journal of Sichuan Forestry Science and Technology, 2011, 32(4):96-98

67.   Li FF, Li ZB, Pan KW*, et al. Research on the improvement effect of the main afforestation tree or bamboo species on physical properties of soil moisture after converting cropland to forest in Chengdu. Journal of Sichuan Forestry Science and Technology, 2011, 32(3)66-68

68.   Zhang XP, Pan KW*, Wang JC, Chen QB. Effects of decomposition of mixed leaf litters of the Castanopsis platyacantha Schima sinensis forest on soil organic carbon. Acta Ecologica Sinica, 2011, 31(6):1582-1593

69.   Mu RPan KW*Wang JCChen QB. Effects of ferulic acidp-hydroxybenzoic acid and their mixture on mineral nitrogen and relative microbial function groups in forest soils. Acta Ecologica Sinica, 2011, 31(3):0793-0800

70.    Luo XY, Pan KW*. A Study of problems and solutions in forest management and conservation in the hilly region of middle Sichuan. Journal of Sichuan Forestry Science and Technology, 2010, 31(4):117-119

71.   Huang R, Pan KW*, Wang JC, Li W. Comparison of soil nitrogen status among three stand types in the semiarid areas of upper reaches of the Minjiang River. Acta Ecologica Sinica201030(5):1210~1216

72.   Gong PM, Pan KW*,Wang JC, et al. Effects of two forms of mineral nitrogenous fertilizers on nematode community composition and diversity in rhizosphere of cucumber in greenhouses. Acta Ecologica Sinica, 2009, 29(2):1009-1016

73.   Liu ZW, Duan EJ, Pan KW, et al. C and N allocation patterns in planted forests and their release patterns during leaf litter decomposition in subalpine area of west Sichuan. Chinese Journal of App lied Ecology, 2009, 28(1):1-6

74.   Li XY, Zhang YB, Pan KW*, et al. Effects of elevated temperature on reproductive phenology and growth of Allium xichuanense and Anemone rivularis in timberline ecotone. Chinese Journal of Ecology, 2009, 28(1):12-18

75.   Zhou XM, Pan KW*, Wang JC. Effects of native Pinus tabulaeformis and exotic Pinus radiata afforestation on soil phosphorus in the upper reaches of Minjiang River. Acta Ecologica Sinica, 2009, 29(12):6630-6637

76.   Zhang JM, Su YM, Pan KW*, et al. A study of the method of determining the area of forest land requisition for the building of the transmission line. Journal of Sichuan Forestry Science and Technology, 2009, 30(6):124-125

77.   Liang XL, Pan KW*, Wang JC. Releasing dynamics of phenolic acid during Zanthoxylum bungeanum litter decomposition and effects of its aqueous extract on soil chemical properties. Acta Ecologica Sinica, 2008, 28(10): 4676-4684

78.   Liu ZW, Gao WJ, Pan KW, et al. Effects of soil mixing incubation of 4 species litters of artificial forests on the biochemical characteristics. Journal of Northwest A & F University, 2007, 7: 48-54

79.   Li X, Pan KW, Gao P, et al. Effect of the pesticide on soil microbe and enzyme activity. Journal of Anhui Agri. Sci, 2007, 21: 6510-6512

80.   Chen J, Pan KW, Bi SR, Gu B, Wang JC, Liu XR. Study on stem apex tissue culture of Zingiber officinale and the control of contamination rate. Journal of Changjiang Vegetables, 2007, 1: 38-40

81.   Liu ZW, Gao WJ, Pan KW, et al. Influences of soil mixing of different forests on the biochemical characteristics and litter decomposition on upper reach of Minjiang River. Acta Ecologica Sinica, 2007, 27(10):4149-4156

82.   Chen J, Pan KW, Gu B. Physiological function and mechanism of abscisic acid in plants under stress. Plant Physiology Communications, 2006, 42(6):1176-1182

83.   Chen J, Pan KW, Gu B, et al. Atractylodes macrocephala rapid propagation by direct shoot and plant regeneration by leaf. Chinese Agricultural Science Bulletin, 2006, 22(11):65-69

84.   Song L, Pan KW*, Wang JC, Ma YH. Effects of phenolic acids on seed germination and seedling antioxidant enzyme activity of alfalfa. Acta Ecologica Sinica, 20062610):3393-4303

85.   Lu K, Pan KW*, Wang JC. Effects of Zanthoxylum bungeanum leaf extract on soil microbe quantity and enzyme activities. Chinese Journal of Applied Ecology, 2006179):1649-1654

86.   Liu ZW, Pan KW, Du HX, et al. Effects of C and N disturbances on decomposition of leaf litter in forests. Journal of Soil and Water Conservation, 2006, 20(4):160-162

87.   Du HX, Liu ZW, Pan KW, et al. Effects of external source C, N disturbances on enzymes activities of forest soil. Journal of Northwest Forestry University, 2006, 21(2):35-38

88.   Liu ZW, Pan KW, Du HX. Research progress on nitrogen relationship between forest plants, litter and soil microorganisms. Journal of Northwest Forestry University200621(1)72-75

89.   Liu ZW, Gao WJ, Pan KW. Discussion on the study methods and models of litter decomposition. Acta Ecologica Sinica, 2006, 26(6):1993-2000

90.   Wang JC, Pan KW*,Wu N. The study on varietal differences in allelopathic potential of Chinese prickly ash. Acta Ecologica Sinica, 2005, 25(7):77-85.

91.   Chen YP, Pan KW*, Wu N, et al. Effect of litter quality and decomposition on mineralization in soil of Castanopsis platycantha-Schima sinensis forest. Chin J Appl Environ Biol, 2005,11(2)146-151

92.   Wang JC, Pan KW*,Wu N, et al. Effect of simplied agroforest system of Sichuan pepper on some ecological factors. Chin J Appl Environ Biol, 2005, 11(1)36-39

93.   Liu ZW, Pan KW. Problems in Olson's litter decomposition model and its revision. Journal of Northwest Sci-Tech Univ. of Agri. and For. (Nat. Sci. Ed.), 2005, 33(1):69-70

94.   Gao XBLiu ZWPan KW, et al. Preliminary studies on soil enzymes activities of typical forest ecosystem in Minjiang Areas. Journal of Northwest Forestry University2005203):1-5

95.   Li FH,Wang YT, Pan KW. Progress, problems and prospects of maize Curvularia leaf spot disease. Plant Protection, 2004, 30(6):5-10

96.   Wang JC, Pan KW*, Li FH. Status and prospects of allelopathy in molecular level and soil ecosystem. Chinese Journal of Ecology, 2004, 23(6):125-130

97.   Pan KW, Wu N, Pan KZ. A discussion on the issues of the re-construction of ecological shelter zone on the upper reaches of the Yangtze River. Acta Ecologica Sinica, 2004, 24(3)617-629

98.   Pan KW, He J and Wu N. Effect of forest litter on microenvironment conditions of forestland. Chinese Journal of Applied Ecology, 2004, 15(1):153-158

99.   Zhao CM, Chen QH, Qiao YK, Pan KW. Structure and spatial pattern of a nature Abies faxoniana population on the eastern edge of Qinghai Tibetan Plateau. Acta Phytoecological Sinica, 2004, 28(3):341-350

100.Liu ZW, Pan KWLi YS. Discussion on nutrients relationship between the primary producers and decomposers in forest-ecosystem. Science of Soil and Water Conservation2003(4)102-105

101.Pan KW, Zhang YB. Percentage of soil absolute moisture content at different succession stages of Castanopsis platyacantha Schima sinensis formation in central subtropical zone of Sichuan Province. Chinese Journal of Applied Ecology, 2003, 14(2):165-169

102.Zhang YBWang KYHu TXPan KW, et al. Plant species diversity of understory communities in Castanopsis platycantha-Schima sinensis formation. Chin J Appl Environ Biol, 2003, 9(5):465-470

103.Zhang YBWang KYHu TXPan KW, et al. Analysis on growth of Castanopsis platycantha in two different stands. Chin J Appl Environ Biol, 2003, 9(4):336-340

104.Zhang YM, He J, Pan KW*, et al. Potential contribution of the soil seed banks to the restoration of native plants. Chin J Appl Environ Biol, 2003, 9(3):326-332

105.WanT, Zhang JM, Pan KW. Ecological function and restoration strategies of forests in the regions of evergreen broad leaved forest in the lower and middle mountain areas in the Upper Reaches of the Yangtze River. Journal of Sichuan Forestry Science and Technology, 2003, 24(1):56-60

106.Wan T, Zhang JM, Pan KW. The growth law of the mean individual from artificial community of 10 years old Cercidiphyllum japonicum. Journal of Sichuan Forestry Science and Technology, 2002, 3:34-38

107.Zhao CM, Chen QH, Qiao YK, Pan KW. Dynamics of species diversity in the restoration process of artificial spruce conifer forest in the eastern edge of Qinghai Tibetan Plateau. Acta Phytoecological Sinica, 2002, 26:20-29

108.Pan KW, Zhang YM, Liu ZG,et al. Comparative study on microclimate of vegetation over different successional stages of Castanopsis platycantha-Schima sinensis formation the central subtropical zone in Sichuan province. Acta Phytoecological Sinica, 2002, 26(2):195-202

109.Liu Q, Qiao YK, Wu N, Pan KW, et al. Mechanism of ecological degradation of mountain ecosystem in the upper reaches of the Minjiang River: effects of keystone populations. Resources and Environment in the Yangtze Basin, 2002, 11(3):274-278

110.Pan KW and Liu ZG. Grey correlation and cluster analysis on relationship between Cercidiphyllum japonicum community and its environment. Chinese Journal of Applied Ecology, 200112(2):161-167

111.Wu Y, Liu Q, Qiao YK, Pan KW, et al. Species diversity changes in subalpine coniferous forests of different restoration stages and their effects on soil properties. Acta Phytoecological Sinica, 2001, 25(6):648-655

112.Pan KW, Liu ZG. The distribution and circulation of nutrient element contents and accumulation in the young artificial Cercidphyllum japonicum community. Scientia Silvae Sinicae, 2001, 37(2):1-12

113.Liu ZGPan KW. Difficulties and strategies for Grain for Green Project in the upper reaches of the Yangtze River. Resources and Environment in the Yangtze Basin, 2001, 10(5): 426-431

114.Pan KW, Liu CL, Liu ZG, et al. Climate resource and the utilization strategies in Mt. Wawushan area. Journal of Mountain Science, 2000, 18(2):1-8

115.Pan KW, Liu ZG. Biomass from artificial community of 10 a Cercidiphyllum japonicum. Chin J Appl Environ Biol, 1999, 5(2): 121-130.

116.Pan KW. Structures of the artificial mixed forests in the clear-cut area of subalpine dark coniferous forests on the upper branches of the Minjing River. Journal of Wuhan Botanical Research, 1999, 7(2)130-136

117.Pan KW. The content of available Al of the top soil and its effect on the growth of Cercidiphyllum japonicum communities in Dagou valley in Sichuan. Journal of Mountain Science, 1999, 17(2)147-151

118.Pan KW, Yang DS, Jiang X. The change and change trend prediction of forest soil erosion after improvement of the deteriorated forests of masson pine (Pinus massoniana) in Sichuan Basin. Journal of Soil Erosion and Soil and Water Conservation, 1995, 1(1): 48-53

119.Pan KW, Yang DS, Jiang X. On the technology of multiple-stratum management system with the deteriorated forests of masson pine (Pinus massonia). Journal of Sichuan Agricultural University, 1995, 13(2): 188-191, 149


Jiang HB, Pan KW, Li N, Zhou XM and Chen JZ. Nondestructive detection method for three weft structure of plant root system201510028033.1


Selected Projects

2019-2022: National Natural Science Foundation of China “Driving Mechanism of Forest Soil Micro-food Webs on β-Glucosidase Microbial Functional Groups and Its Response to Warming and Drought: A Comparative Study of Spruce Contrasting Forests in Poland and China”

2020-2023: National Natural Science Foundation of China “Strategic regulation of hydraulic conductance of Phoebe zhennan seedling by nitrogen and phosphorus supplementation under drought stress: Towards unraveling the mechanisms of morphological, physiological and anatomical changes of root branching order”

2016-2020: Sub-task of National Key Research and Development Plan:“Formation mechanism of ecological pattern in Southwestern China”.

2015-2019: Sub-project of National Science and Technology Support Program: “Management techniques of resource-efficient agroforestry in the low and hilly land in the southwestern China”.

2016-2017: Large enterprises entrusted project: "Ecological impact assessment of construction of the Cosmic Ray Observation Station in High Altitude Area on Haizishan National Nature Reserve in Sichuan".

2016-2017: STS project of the Chinese Academy of Sciences: "technology integration and demonstration of agroforestry system".

2016-2018: National Natural Science Foundation of China Youth Foud:“Response of soil nematode micro food web to nitrogen deposition and extreme drought.

2014-2017: National Natural Science Foundation of China: “The structure and function of soil food webs and its responses of drought and enhanced rainfall between the comparable systems of Zanthoxylum bungeanum monoculture and mixtures with cover crops on the upper reaches of Minjiang River”.

2012-2014: National Natural Science Foundation of China: “The effect of simulated warming and nitrogen deposition on the ecological stoichiometry relationship of C, N and P in shrub plant— soil of alpine timberline ecotone”.

2011-2013: National Natural Science Foundation of China: “The response of respiratory metabolism and mitochondrial defense pathways of reactive oxygen species in Fargesia root to drought stress and phosphorus addition.”

2011-2015Sub-task of National Science and Technology Support Program: “The control technology of valley soil erosion.”

2010-2012: National Natural Science Foundation of China Youth Foud : “Dynamics of soil dissolved organic and nitrogen and their responses to litter decomposition under the typical artificial plantations of the upper reaches of Minjiang River”.

2006-2010: Sub-project of National Key Technologies R & D Program of China: “Techniques and demonstration of ecological planting and breeding as well as agroforestry in the surrounding regions of natural heritages sites.”

2008: Global Environment Fund Programme: “The biodiversity surveys and risk assessment of Wenchuan disaster regions in Sichuan Province”.

2005-2009: Sub-project of National Key Technologies R & D Program of China: “Configuration of eco-agricultural model as well as the demonstration of its regulated and controlled technologies in the valley”.

1998-2000: Sub-task of Key projects of Chinese Academy of Sciences "Natural restoration of degraded evergreen broad leaved forest in mid subtropical zone of China"